4.5.4 Arrested succession
Fringe wetlands along rivers rarely pass beyond an early successional stage, because floods flush out accumulated silt and detritus. Similarly, flood waters moving rapidly over flood wetlands shift sediment around and break down vegetation. Rivers in areas flat enough to develop aquatic marginal wetlands will also tend to change their course frequently, either by gradual erosion or cat- astrophically in flash floods, unless artificially constrained, and will cut through wetland areas. For these reasons, wetlands along river floodplains can be very heterogeneous, but rarely progress to terrestrial or mire stages.
If flood waters rise slowly with little kinetic energy, then physical changes are less likely to occur; encroachment by terrestrial species will be suppressed but other successional processes are unaffected. Succession will remain in an arrested state, however, if the dry season allows the mineralisation of organic layers that have built up during the flood. If a river's ability to flood is curtailed, by damming or embanking, then terrestrial vegetation will quickly dominate, because the wetland has been cut off from its supply of water.
Fringing marine wetlands may go through cyclical changes in which succession proceeds to a given stage and then is reset by storm activity. Dry land stages will only be reached rapidly if the wetland is actively expanding, for example through rapid siltation.