A rather sophisticated 'mutual-aid' style variation on the monogamous family unit
is found in the
tamarins and marmosets: all members of the family, from father to mature offspring, take part in the
care of the newborn young. Another interesting slant to the lives of the callitrichids is their heavy
reliance on scent marking to define territory, a custom which is rare or absent in most Old and New
World monkeys but common among the Madagascan lemurs. Saddle-back tamarins apply scent
marks to lianas and slim branches using three different methods: ano-genital, by sitting upright and
rubbing their backsides against the branch with a rotating action, possibly at the same time
reinforcing the message with a spattering of urine; suprapubic marking, in which the animal lies
prone and pulls itself forwards with its arms, massaging the suprapubic region against the bark to
smear it with scent; and, much more rarely, sternal marking, in which the chest is rocked to and
fro against the bark. As in lemurs, most of this activity is concentrated around the periphery of the
home range. Scent marking may be applied more intensively on a number of favourite spots, such
as a certain liana which may attract a thorough going over from the whole group in concert.