4.4.3 Species diversity
Species diversity in wetlands is determined to a large extent by succession and seasonal change, both of which allow a wide range of species assemblages to coexist in close proximity. Some wetland types can be species-poor, good examples being the large tracts of swamp dominated by a single species, such as reed (Phragmites australis) or papyrus (Cyperus papyrus), lacking ground vegetation because they are permanently inundated, and impeding submerged vegetation through shading and the large volume of decomposing detritus. A feature of wetland areas, however, is that so often several different habitat types, each with its representative species, are mixed together in a small area. The precise adaptations of different wetland plants to specific degrees of waterlogging contribute to this high species diversity, with very small changes in topography across a small area allowing different species assemblages to coexist. An embanked river, for example, has an open water habitat, a fringing wetland, a normally dry levee and a flood wetland, all within a few metres of each other.
Just as the water table produces spatial patterns in wetland biota, so any fluctuations will create temporal patterns in species diversify.  In any one season. diversity may be low, but when all seasons are considered together, diversity is enhanced by seasonal turnover of species.